This function allows to test fourth-corner statistics using constrained null
models (for traits and/or environmental variables). If the argument
phyloORorthobasis
is specified, random traits are
phylogenetically-constrained to preserve the global autocorrelation (Moran's
I) and the phylogenetic structures at multiple scales. If not, standard
permutations are used. If the argument listwORorthobasis
is specified,
random environmental variables are spatially-constrained to preserve the
global autocorrelation (Moran's I) and the spatial structures at multiple
scales. If not, standard permutations are used. Multiscale property is
defined by the power spectrum (i.e. decomposition of the variance of the
original variables) on a basis of orthonormal eigenvectors (Moran's
Eigenvector Maps, MEM).
An object generated by the fourthcorner
function.
an object of the class listw
(spatial
weights) created by the functions of the spdep package or an object
of class orthobasis
an object of the class phylo
(phylogeny)
created by the functions of the ape package or an object of class
orthobasis
generated by functions of adephylo
(me.phylo
)
an integer
indicating the number of replicates
an character specifying which algorithm should be used to
produce spatial replicates (see msr.default
).
further arguments of the msr.default
function.
An object of class 4thcorner
randomized replicates.
Braga, J., Thuiller, W., ter Braak, C.J.F. and Dray, S. (2018) Integrating spatial and phylogenetic information in the fourth-corner analysis to test trait-environment relationships. Ecology, 99:2667-2674.
if(require("ade4", quietly = TRUE) & require("adephylo", quietly = TRUE)
& require("spdep", quietly = TRUE) & require("ape", quietly = TRUE)){
data(mafragh, package = "ade4")
fr1 <- fourthcorner(mafragh$env, mafragh$flo, mafragh$traits$tabQuantitative, nrepet = 49)
phy <- read.tree(text = mafragh$tre)
lw <- nb2listw(mafragh$nb)
fr1.msr <- msr(fr1, listwORorthobasis = lw, phyloORorthobasis = phy)
fr1
fr1.msr
}
#>
#> Attaching package: ‘ape’
#> The following object is masked from ‘package:adegraphics’:
#>
#> zoom
#> Fourth-corner Statistics
#> ------------------------
#> Permutation method msr ( 49 permutations)
#>
#> Adjustment method for multiple comparisons: holm
#> call: msr.4thcorner(x = fr1, listwORorthobasis = lw, phyloORorthobasis = phy)
#>
#> ---
#>
#> Test Stat Obs Std.Obs Alter Pvalue
#> 1 Clay / Lfp r 0.014241856 0.31344983 two-sided 0.8
#> 2 Silt / Lfp r -0.076256133 -1.04348067 two-sided 0.38
#> 3 Sand / Lfp r 0.063614861 0.83723555 two-sided 0.4
#> 4 K2O / Lfp r 0.031979355 0.55194697 two-sided 0.78
#> 5 Mg++ / Lfp r 0.014982765 0.30489925 two-sided 0.86
#> 6 Na+/100g / Lfp r 0.093499930 1.58354671 two-sided 0.14
#> 7 K+ / Lfp r 0.149968927 1.66354996 two-sided 0.04
#> 8 Conductivity / Lfp r 0.080497038 1.39839433 two-sided 0.18
#> 9 Retention / Lfp r -0.067943687 -0.78196411 two-sided 0.5
#> 10 Na+/l / Lfp r 0.063272819 1.23379224 two-sided 0.2
#> 11 Elevation / Lfp r -0.129537428 -1.42118199 two-sided 0.2
#> 12 Clay / Min height r 0.061180871 1.24718751 two-sided 0.2
#> 13 Silt / Min height r -0.052382325 -0.97141557 two-sided 0.36
#> 14 Sand / Min height r -0.027907397 -0.70638146 two-sided 0.46
#> 15 K2O / Min height r 0.047550545 0.86529722 two-sided 0.44
#> 16 Mg++ / Min height r -0.023023055 -0.21889128 two-sided 0.96
#> 17 Na+/100g / Min height r 0.027191299 0.68785365 two-sided 0.5
#> 18 K+ / Min height r 0.095029793 1.75996060 two-sided 0.06
#> 19 Conductivity / Min height r -0.018899856 -0.27560449 two-sided 0.8
#> 20 Retention / Min height r 0.004165428 0.15953651 two-sided 0.96
#> 21 Na+/l / Min height r -0.046119268 -0.77623227 two-sided 0.46
#> 22 Elevation / Min height r 0.061211127 0.75806348 two-sided 0.38
#> 23 Clay / Max height r 0.010762151 0.21379340 two-sided 0.82
#> 24 Silt / Max height r -0.030835126 -0.45487793 two-sided 0.72
#> 25 Sand / Max height r 0.027229507 0.47114234 two-sided 0.58
#> 26 K2O / Max height r 0.063923177 1.21898682 two-sided 0.24
#> 27 Mg++ / Max height r 0.010886210 0.25253410 two-sided 0.78
#> 28 Na+/100g / Max height r 0.026525395 0.63579493 two-sided 0.48
#> 29 K+ / Max height r 0.109401849 1.49608195 two-sided 0.18
#> 30 Conductivity / Max height r 0.002446542 0.04059237 two-sided 0.98
#> 31 Retention / Max height r -0.005456736 0.06726077 two-sided 0.98
#> 32 Na+/l / Max height r -0.018577793 -0.47742690 two-sided 0.68
#> 33 Elevation / Max height r 0.033495574 0.58742707 two-sided 0.56
#> Pvalue.adj
#> 1 1
#> 2 1
#> 3 1
#> 4 1
#> 5 1
#> 6 1
#> 7 1
#> 8 1
#> 9 1
#> 10 1
#> 11 1
#> 12 1
#> 13 1
#> 14 1
#> 15 1
#> 16 1
#> 17 1
#> 18 1
#> 19 1
#> 20 1
#> 21 1
#> 22 1
#> 23 1
#> 24 1
#> 25 1
#> 26 1
#> 27 1
#> 28 1
#> 29 1
#> 30 1
#> 31 1
#> 32 1
#> 33 1
#>
#> ---
#> Signif. codes: 0 ‘***’ 0.001 ‘**’ 0.01 ‘*’ 0.05 ‘.’ 0.1 ‘ ’ 1